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Cold Spr. Harb. Symp. quant. Biol. 20, 33. KOJIMA, K. (1959). Proc. nat. Acad. Sci. 45,989. KOLMOGOROV, A. (1931). Math. Ann. 104, 415. LEVINS, R. (1961). General Systems, 6, 33. LEVINS; R. (1962). Amer. Nat. 96, (891), 361. LEVINS. R. (1963). Amer. Nat. 97. (893). 15. MORAN, P. k. P:(1958). Hum. Giiet. 53, 1. MORAN, P. A. P. (1959). J. Aust. math. Sot. 1, 121. MORAN, P. A. P. (1960). Trans. Camb. phiI. Sot. 57, 304. MORAN, P. A. P. (1961). UHLENBECK, G. E. & ORNSTEIN, L. S. WRIGHT, S. (1931). Genetics, 16, 97.

20: 276-286. Da Cunha, A. , and Th. Dobzhansky, 1954, A furtherstudy of chromosomal polymorphism in Drosophila willistoni in relation to its environment. Evolution 8: 119-134. , 1952, The varieties of evolutionary opportunity. Rev. Canad. Biol. 11: 305-388. , 1953, Genetic equilibrium when more than one ecological niche is available. Am. Naturalist 87: 311-???. , Th. Dobzhansky and 0. Pavlovsky, 1954, Interaction of adaptive values in polymorphic experimental populations of DrosophZla pseudoobscura.

20: 276-286. Da Cunha, A. , and Th. Dobzhansky, 1954, A furtherstudy of chromosomal polymorphism in Drosophila willistoni in relation to its environment. Evolution 8: 119-134. , 1952, The varieties of evolutionary opportunity. Rev. Canad. Biol. 11: 305-388. , 1953, Genetic equilibrium when more than one ecological niche is available. Am. Naturalist 87: 311-???. , Th. Dobzhansky and 0. Pavlovsky, 1954, Interaction of adaptive values in polymorphic experimental populations of DrosophZla pseudoobscura.

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Theory of Fitness in a Heterogeneous Environment by Richard Levins


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